Prof John Harwood

Prof John Harwood


Population and Conservation Biology : Effects of individual variation and spatial structure on the population dynamics, genetics and epidemiology of vertebrates, particularly marine mammals. Interactions between anthropogenic activities and ecosystem function.

Research groups:


NERC Sea Mammal Research Unit

source: symbiosis


Recent publications listed in research@st-andrews
Tyack, PL, Bailey, H, Crocker, D, Estes, JE, Francis, CD, Harwood, J, Schwacke, L, Thomas, LJ & Wartzok, D 2016, Approaches to Understanding the Cumulative Effects of Stressors on Marine Mammals. National Academies Press, Washington DC. National Academies Press, Washington DC. DOI: 10.17226/23479
Fleishman, E, Costa, DP, Harwood, J, Kraus, S, Moretti, D, New, LF, Schick, RS, Schwarz, LK, Simmons, SE, Thomas, L & Wells, RS 2016, 'Monitoring population-level responses of marine mammals to human activities' Marine Mammal Science, vol 32, no. 3, pp. 1004-1021. DOI: 10.1111/mms.12310
New, LF, Clark, J, Costa, D, Fleishman, E, Hindell, M, Klanjšček, T, Lusseau, D, Kraus, S, McMahon, C, Robinson, P, Schick, RS, Schwartz, L, Simmons, S, Thomas, L, Tyack, PL & Harwood, J 2014, 'Using short-term measures of behaviour to estimate long-term fitness of southern elephant seals.' Marine Ecology Progress Series, vol 496, pp. 99-108. DOI: 10.3354/meps10547
Schick, RS, New, L, Thomas, L, Costa, D, Hindell, M, McMahon, C, Robinson, P, Simmons, S, Thums, M, Harwood, J & Clark, J 2013, 'Estimating resource acquisition and at-sea body condition of a marine predator' Journal of Animal Ecology, vol 82, no. 6, pp. 1300-1315. DOI: 10.1111/1365-2656.12102
Smout, SC, Asseburg, C, Matthiopoulos, J, Fernández, C, Redpath, S, Thirgood, S & Harwood, J 2010, 'The functional response of a generalist predator' PLoS One, vol 5, no. 5, e10761. DOI: 10.1371/journal.pone.0010761
Hall, AJ & Harwood, J 2009, Mass Mortalities. in WF Perrin, B Wursig & JGM Thewissen (eds), Encyclopedia of Marine Mammals. Academic Press Inc., pp. 709-711.
Cabrelli, A, Harwood, J, Matthiopoulos, J, New, LF & Thomas, L 2009, A critical review of the literature on population modelling. CREEM Technical Report, no. 2009-2, University of St Andrews.
Mollett, A, Schofield, C, Miller, I, Harwood, J, Harris, CM & Donovan, CR 2009, Environmental Risk Management Capability: Advice on Minimising the Impact of Both Sonar and Seismic Offshore Operations on Marine Mammals. in Offshore Europe Oil and Gas Conference and Exhibition : proceedings : 8-11 September 2009, Aberdeen Exhibition and Conference Center UK, Aberdeen UK. Society of Petroleum Engineers, pp. 1-8, 2009 SPE Offshore Europe Oil and Gas Conference and Exhibition, Aberdeen, United Kingdom, 8 September. DOI: 10.2118/124301-MS
Matthiopoulos, J, Graham, K, Smout, SC, Asseburg, C, Redpath, S, Thirgood, S, Hudson, P & Harwood, J 2007, 'Sensitivity to assumptions in models of generalist predation on a cyclic prey' Ecology, vol 88, no. 10, pp. 2576-2586. DOI: 10.1890/06-0483.1
Asseburg, C, Harwood, J, Matthiopoulos, J & Smout, SC 2006, The functional response of generalist predators, and their implications for the monitoring of marine ecosystems. in IL Boyd, S Wanless & CJ Camphuysen (eds), Top predators in marine ecosystems: their role in monitoring and management . Cambridge University Press, pp. 262-274.
Gaggiotti, OE, Jones, F, Lee, WM, Amos, W, Harwood, J & Nichols, RA 2002, 'Patterns of colonization in a metapopulation of grey seals' Nature, vol 416, no. 6879, pp. 424-427. DOI: 10.1038/416424a
Grenfell, BT, Amos, W, Arneberg, P, Bjornstadt, ON, Greenman, JV, Harwood, J, Lanfranchi, P, McLean, A, Norman, RA, Read, AF & Skorping, A Visions for future research in wildlife epidemiology.


The Secretary
Scottish Oceans Institute
Institiud Chuantan na h-Alba
Gatty Marine Laboratory
East Sands St Andrews
KY16 8LB

tel: +44 (0) 1334 463472
fax: +44 (0) 1334 463443