Prof John Harwood

Prof John Harwood


Population and Conservation Biology : Effects of individual variation and spatial structure on the population dynamics, genetics and epidemiology of vertebrates, particularly marine mammals. Interactions between anthropogenic activities and ecosystem function.

Research groups:


NERC Sea Mammal Research Unit

source: symbiosis


Research Overview:

Population and Conservation Biology

Effects of individual variation and spatial structure on the population dynamics, genetics and epidemiology of vertebrates, particularly marine mammals. Interactions between anthropogenic activities and ecosystem function.



50 (of 83 /dk/atira/pure/researchoutput/status/published available) for jh17 (source: University of St Andrews PURE)
Please click title of any item for full details

Context-dependent variability in the predicted daily energetic costs of disturbance for blue whales Enrico Pirotta, David E Cade, John Calambokidis, Daniel P Costa, James A Fahlbusch, Ari S Friedlaender, Jeremy A Goldbogen, John Harwood, Elliott L Hazen, Leslie New, Brandon L Southall
Conservation Physiology 2021 vol. 9
A decision framework to identify populations that are most vulnerable to the population level effects of disturbance John Harwood, Ruth Joy, Catriona M Harris
Conservation Science and Practice 2020 vol. 2
Expenditure, investment, and acquisition of energy in southern elephant seals Mike Fedak, Tom A. Arnbom, Bernie J McConnell, C. Chambers, Ian Boyd, John Harwood, T. S. McCann
2020 pp. 354-373
Methods for monitoring for the population consequences of disturbance in marine mammals Rachael R. Sinclair, John Harwood
Frontiers in Marine Science 2020 vol. 7
Propensity for risk in reproductive strategy affects susceptibility to anthropogenic disturbance E. Pirotta, Vincent Hin, Marc Mangel, Leslie New, Daniel P. Costa, André M. de Roos, John Harwood
American Naturalist 2020 vol. 196 pp. E71–E87
The Epidemiological Framework for Biological Invasions (EFBI) Philip E. Hulme, Robert Freckleton, Rosemary S. Hails, Matt Hartley, John Harwood, Glenn Marion, Graham Smith, Mark Williamson
NeoBiota 2020 vol. 62 pp. 161-192
Anthropogenic disturbance in a changing environment Enrico Pirotta, Marc Mangel, Daniel P. Costa, Jeremy Goldbogen, John Harwood, Vincent Hin, Ladd M. Irvine, Bruce R. Mate, Elizabeth A. McHuron, Daniel M. Palacios, Lisa K. Schwarz, Leslie Frances New
Oikos 2019 vol. 128 pp. 1340-1357
Bio-energetic modeling of medium-sized cetaceans shows high sensitivity to disturbance in seasons of low resource supply Vincent Hin, John Harwood, André M. de Roos
Ecological Applications 2019 vol. 29
Modelling the population size and dynamics of the British grey seal Len Thomas, Debbie JF Russell, Callan David Duck, Chris Morris, Mike Lonergan, David Thompson, John Harwood
Aquatic Conservation: Marine and Freshwater Ecosystems 2019 vol. 29 pp. 6-23
Population-level consequences of seismic surveys on fishes Hans Slabbekoorn, John Dalen, Dick de Haan, Hendrik V. Winter, Craig Radford, Michael A. Ainslie, Kevin D. Heaney, Tobias van Kooten, Len Thomas, John Harwood
Fish and Fisheries 2019 vol. 20 pp. 653-685
A simulation approach to assessing environmental risk of sound exposure to marine mammals Carl Robert Donovan, Catriona M Harris, Lorenzo Milazzo, John Harwood, Laura Helen Marshall,
Ecology and Evolution 2017 vol. 7 pp. 2101-2111
Approaches to Understanding the Cumulative Effects of Stressors on Marine Mammals Peter Lloyd Tyack, Helen Bailey, Daniel Crocker, James E Estes, Clinton D Francis, John Harwood, Lori Schwacke, Len Thomas, Douglas Wartzok
Monitoring population-level responses of marine mammals to human activities Erica Fleishman, Daniel P. Costa, John Harwood, Scott Kraus, David Moretti, Leslie F. New, Robert Schilling Schick, Lisa K. Schwarz, Samantha E. Simmons, Len Thomas, Randall S. Wells
Marine Mammal Science 2016 vol. 32 pp. 1004-1021
Understanding the population consequences of acoustic disturbance for marine mammals John Harwood, Carl Robert Donovan, Robert Schilling Schick, Len Thomas, Leslie Frances New
2016 pp. 417-423
An interim framework for assessing the population consequences of disturbance Robert Schilling Schick, Carl Robert Donovan, Mark Burgman, Len Thomas, John Harwood
Methods in Ecology and Evolution 2015 vol. 6 pp. 1150-1158
Modelling production per unit of food consumed in fish populations Mauricio A. Barrientos, Andres C. Milessi, J. C. Quiroz, John Harwood
Journal of Theoretical Biology 2015 vol. 365 pp. 67-75
Predicting the effects of human developments on individual dolphins to understand potential long-term population consequences Enrico Pirotta, John Harwood, Paul Thompson, Leslie New, Barbara Cheney, Monica Arso Civil, Philip Steven Hammond, Carl Robert Donovan, David Lusseau
Proceedings of the Royal Society B: Biological Sciences 2015 vol. 282
A risk function for behavioral disruption of Blainville’s beaked whales (Mesoplodon densirostris) from mid-frequency active sonar Len Thomas, Tiago A. Marques, John Harwood, Ashley Dilley, Bert Neales, Jessica Shaffer, E Mccarthy, Leslie Frances New, S Jarvis, Ron Morrissey
PLoS One 2014 vol. 9
Activities, motivations and disturbance Enrico Pirotta, Leslie Frances New, John Harwood, David Lusseau
Ecological Modelling 2014 vol. 282 pp. 44-58
Errutum Len Thomas, Tiago A. Marques, John Harwood, A. Dilley
PLoS One 2014 vol. 9
Modelling prey consumption and switching by UK grey seals Sophie Caroline Smout, Anna Rindorf, Philip Steven Hammond, John Harwood, Jason Matthiopoulos
ICES Journal of Marine Science 2014 vol. 71 pp. 81-89
Using short-term measures of behaviour to estimate long-term fitness of southern elephant seals. Leslie Frances New, James Clark, Daniel Costa, Erica Fleishman, Mark Hindell, Tin Klanjšček, David Lusseau, Scott Kraus, Clive McMahon, Patrick Robinson, Robert Schilling Schick, Lisa Schwartz, Samantha Simmons, Len Thomas, Peter Lloyd Tyack, John Harwood
Marine Ecology Progress Series 2014 vol. 496 pp. 99-108
Applying distance sampling to fin whale calls recorded by single seismic instruments in the northeast Atlantic. Danielle Harris, Luis Matias, Len Thomas, John Harwood, Wolfram Geissler
Journal of the Acoustical Society of America 2013 vol. 134
Estimating resource acquisition and at-sea body condition of a marine predator Robert Schilling Schick, Leslie New, Len Thomas, Daniel Costa, Mark Hindell, Clive McMahon, Patrick Robinson, Samantha Simmons, Michele Thums, John Harwood, James Clark
Journal of Animal Ecology 2013 vol. 82 pp. 1300-1315
Modelling the biological significance of behavioural change in coastal bottlenose dolphins in response to disturbance Leslie Frances New, John Harwood, Len Thomas, Carl Robert Donovan, James Clark, Gordon Drummond Hastie, Paul Thompson, Barbara Cheney, Lindesay Alexandra Sarah Scott-Hayward, David Lusseau
Functional Ecology 2013 vol. 27 pp. 314-322
Uncovering the links between foraging and breeding regions in a highly mobile mammal Debbie JF Russell, Bernie J McConnell, David Thompson, Callan David Duck, Chris Morris, John Harwood, Jason Matthiopoulos
Journal of Applied Ecology 2013 vol. 50 pp. 499-509
A simulation-based method for quantifying and mitigating the effects of anthropogenic sound on marine mammals Carl Robert Donovan, Catriona M Harris, John Harwood, Lorenzo Milazzo
2012 pp. 1551-1556
Geographic variation of the major histocompatibility complex in Eastern Atlantic grey seals (Halichoerus grypus). K Cammen, J.I. Hoffman, L.A. Knapp, John Harwood, W. Amos
Molecular Ecology 2011 vol. 20 pp. 740-752
Comparison of the 1988 and 2002 phocine distemper epizootics in British harbour seal Phoca vitulina populations Mike Lonergan, Ailsa Jane Hall, Hal Thompson, Paul M. Thompson, Patrick Pomeroy, John Harwood
Diseases of Aquatic Organisms 2010 vol. 88 pp. 183-188
The functional response of a generalist predator Sophie Caroline Smout, C Asseburg, Jason Matthiopoulos, Carmen Fernández, S Redpath, S Thirgood, John Harwood
PLoS ONE 2010 vol. 5
A critical review of the literature on population modelling Abigail Cabrelli, John Harwood, Jason Matthiopoulos, Leslie Frances New, Len Thomas
Environmental Risk Management Capability: Advice on Minimising the Impact of Both Sonar and Seismic Offshore Operations on Marine Mammals Alex Mollett, Colin Schofield, Ian Miller, John Harwood, Catriona M Harris, Carl Robert Donovan
2009 pp. 1-8
Integrated Data Analysis in the Presence of Emigration and Mark Loss Ruth King, John Harwood, Morten Frederiksen, Michael P. Harris, Sarah Wanless
Journal of Agricultural, Biological and Environmental Statistics 2009 vol. 14 pp. 411-431
Mass Mortalities Ailsa Jane Hall, John Harwood
2009 pp. 709-711
Getting beneath the surface of marine mammal - fisheries competition Jason Matthiopoulos, Sophie Caroline Smout, Arliss Winship, David Thompson, Ian Boyd, John Harwood
Mammal Review 2008 vol. 38 pp. 167-188
Passive acoustic detection of deep-diving beaked whales WMX Zimmer, John Harwood, Peter Lloyd Tyack, Mark Johnson, Peter T Madsen
Journal of the Acoustical Society of America 2008 vol. 124 pp. 2823-2832
Travelling through a warming world: climate change and migratory species RA Robinson, John Harwood, None and 18 others
Endangered Species Research 2008
Embedding population dynamics models in inference. Stephen Terrence Buckland, Ken B Newman, C Fernandez, Len Thomas, John Harwood
Statistical Science 2007 vol. 22 pp. 44-58
Influence of the physical environment and conspecific aggression on the spatial arrangement of breeding grey seals Catriona M Harris, Jason Matthiopoulos, John Harwood
Ecological Informatics 2007 vol. 2 pp. 308-317
Is there a role for ecologists in an ecosystem approach to the management of marine resources? John Harwood
Aquatic Conservation: Marine and Freshwater Ecosystems 2007 vol. 17 pp. 1-4
Sensitivity to assumptions in models of generalist predation on a cyclic prey Jason Matthiopoulos, K Graham, Sophie Caroline Smout, C Asseburg, S Redpath, S Thirgood, P Hudson, John Harwood
Ecology 2007 vol. 88 pp. 2576-2586
Defining matrix population models and fitting them to data Stephen Terrence Buckland, Ken B Newman, Len Thomas, John Harwood
2006 pp. 15-27
Incorporating movement into models of grey seal population dynamics Phil Harrison, Stephen Terrence Buckland, Len Thomas, R Harris, Patrick Pomeroy, John Harwood
Journal of Animal Ecology 2006 vol. 75 pp. 634-645
The functional response of generalist predators, and their implications for the monitoring of marine ecosystems. C Asseburg, John Harwood, Jason Matthiopoulos, Sophie Caroline Smout
2006 pp. 262-274
A unified framework for modelling wildlife population dynamics Len Thomas, Stephen Terrence Buckland, Ken B Newman, John Harwood
Australian and New Zealand Journal of Statistics 2005 vol. 47 pp. 19-34
Defining biologically relevant management units for seal populations in the UK K Everard, John Harwood, Jason Matthiopoulos, Bernie J McConnell, R Sharples, Catriona M Harris, C Z Warnes
Metapopulation consequences of site fidelity for colonially breeding mammals and birds Jason Matthiopoulos, John Harwood, Len Thomas
Journal of Animal Ecology 2005 vol. 74 pp. 716-727
Combining demographic, environmental and genetic data to test hypotheses about colonisation events in metapopulations Oscar Eduardo Gaggiotti, SP Brooks, W Amos, John Harwood
Molecular Ecology 2004 vol. 13 pp. 811-825


The Secretary
Scottish Oceans Institute
Institiud Chuantan na h-Alba
Gatty Marine Laboratory
East Sands St Andrews
KY16 8LB

tel: +44 (0) 1334 463472
fax: +44 (0) 1334 463443